|A modern alytid European frog, Bombina. JCM|
The excavation site was at Plakias (early Tortonian, MN 9), Crete, Greece. Most of the material recovered was fragmentary and precludes precise taxonomic assignment. However, the herpetofauna of Plakias was diverse and composed at least six different taxa: an alytid frog, a crocodilian, two turtles (a pan-trionychid and a geoemydid) and two squamates (an amphisbaenian and a colubroid snake). The crocodilian material represents the first such fossils described from Greece and furthermore, one of the latest occurrences of this group in Europe. The pan-trionychid and the geoemydid represent the oldest occurrences of these groups in Greece and further add to their scarce Miocene record from this country. The first description of a fossil amphisbaenian from Greece is also provided. The new specimens from Plakias add to our knowledge of the Miocene herpetofaunas of southeastern Europe. The single colubroid snake specimen adds further to the published record of Miocene snakes from Greece, whereas the amphisbaenian vertebra from Plakias represents the first described fossil of this group from the country, suggesting that amphisbaenians had a continuous range in the northern Mediterranean area.
Georgalis, G.L., Villa, A., Vlachos, E. and Delfino, M., 2016. Fossil amphibians and reptiles from Plakias, Crete: A glimpse into the earliest late Miocene herpetofaunas of southeastern Europe. Geobios.